a link to her students blog.
Journal articles with more specific information about this subject are:
Pangle, W. M. & Holekamp, K. E. (2010) Functions of vigilance behavior in a social carnivore, the spotted hyaena, Crocuta crocuta. Animal Behaviour. 80: 257-267.
Pangle, W. M & Holekamp, K. E. (2010) Lethal and non-lethal anthropogenic effects on spotted hyenas in the Masai Mara National Reserve. Journal of Mammalogy. 91:154-164.
Kolowski, J. M. & Holekamp, K. E. (2009) Ecological and anthropogenic influences on space use by spotted hyaenas (Crocuta crocuta). Journal of Zoology, London. 277: 23-36.
Boydston, Erin E., et al. "Altered behaviour in spotted hyenas associated with increased human activity." Animal Conservation 6.3 (2003): 207-219.
photos are property of R. Kappler and L. Jonaitis